Pseudoteloceras, a new stephanoceratid genus (Ammonitida) of the lower Humphriesianum Zone (lower Bajocian, Middle Jurassic) from western Tethys

The new dimorphic genus Pseudoteloceras (type-species: P. crosillense gen. et sp. nov.) is defined within the subfamily Stephanoceratinae. It constitutes the terminal step of an early Bajocian phyletic trend that produces Teloceras-like morphologies throughout the lower Humphriesianum Zone of the western Mediterranean-Caucasian Subrealm, at an older chronological interval and with morpho-structural features that stand apart from those of the best known genus Teloceras of the uppermost Humphriesianum and lower Niortense zones at the early/late Bajocian transition. Three new species are described: P. croisillense, P. maerteni and P. boursicoti. Their biochronostratigraphical ranges are confirmed by the distribution in expanded sections of the Subalpine Basin in Submediterranean Province (Digne area) and the taphonomic analysis of fossil assemblages from the condensed sections of the “Oolithe ferrugineuse de Bayeux” Formation in Calvados, North West European Province. The biochronostratigraphical range of Pseudoteloceras gen. nov. is limited to the Romani Subzone, lower Humphriesianum Zone. P. crosillense gen. et sp. nov. marks a well-defined biohorizon of the upper Romani Subzone in the Digne stratigraphical successions. The extreme degree of the cadiconic morphology in the inner whorls of P. boursicoti gen. et sp. nov., and the biostratigraphical gap separating the first appearance of the genus Teloceras in the upper Humphriesianum Zone, support its erection as a new taxon Pseudoteloceras gen. nov. A fourth species, Pseudoteloceras geometricum (Maubeuge), is interpreted as the earliest species of this phyletic lineage, derived from Stemmatoceras and widely distributed through western Tethys in the lower Humphriesianum Zone.