Thalassinoides in the Middle Miocene succession at Siwa Oasis, northwestern Egypt

At Siwa Oasis, the carbonate-rich Middle Miocene Marmarica Formation includes several firmground discontinuity surfaces, characterized by extensive burrow systems. These surfaces are confined to the boundary between the Oasis and Siwa Escarpment members in addition to several intraformational marine firmground discontinuities at several horizons within these two members. The marine trace fossils from all these discontinuities are represented mainly by the ichnogenus Thalassinoides. In all occurrences, Thalassinoides are characterized by a sharp erosional lower contact and grades upward into fossiliferous argillaceous limestone facies. Three Thalassinoides ichnospecies have been reported, including: Th. horizontalis Myrow, 1995, Th. suevicus (Rieth, 1932) and Th. paradoxicus (Woodward, 1830). They are mostly recorded as hypichnial/endichnial forms at the sole of fossiliferous argillaceous limestone facies. Rare exichnial occurrences are also recorded. Thalassinoides ichnospecies displayed considerable variations in their stratigraphic distribution, in addition to remarkable variations in their complexity and dimension. These variations are mostly reflecting environmental conditions prevailed during the trace-maker life. Paleoenvironmental interpretation of these burrows confirmed a well-oxygenated shallow marine environment with extremely slow or nearly nil rate of sedimentation.