New Eoptychoptera (Insecta: Diptera, Ptychopteridae) from the Lower Cretaceous of Spain

A new species, Eoptychoptera cantabrica sp. nov. is described from the Albian El Soplao amber-bearing deposits based on a single male with an elongated proboscis, unknown among extant members of Ptychopteridae. It represents the youngest record of the species-diverse genus Eoptychoptera. A pupa of Eoptychoptera sp. with a long respiratory horn, typical of extant ptychopterids, is described from Las Hoyas limestones, where adult fossil ptychopterids have yet to be discovered. A key to species of Eoptychoptera, based on wing venation, and a distribution map of Cretaceous Ptychopteridae are provided. The climatic preferences and mouthparts of Mesozoic and recent Ptychopteridae are discussed.