Contribution to the environmental ecology of Cryptogramma crispa (L.) R. Br. ex Hooker in the Alps

We studied the habitat characteristics in a number of Cryptogramma crispa stations throughout the Alps. Our objectives were: (1) to define the environmental ecology of C. crispa in the southern part of its distributional area in Europe; (2) to test the reliability of ecological indicator values of soil reaction, nutrient and dispersion status, light and temperature for this species. The habitats of C. crispa mostly consisted in coarse-grained stable screes (on rock slide deposits, talus cones, talus sheets, stone fields and moraines), within a broad elevation range with maximum frequency in the subalpine and lower alpine vegetation belts. At higher altitudes, C. crispa was mostly found on South-exposed slopes with high potential irradiance, whereas at low altitude it was most abundant on North-exposed rather gentle slopes formed of big boulders. The screes colonized by C. crispa were formed of acid metamorphic rocks or igneous rocks and had a coarse texture. The soil accumulated in the spaces among boulders had sandy texture and acidic reaction, with medium to high carbon and total nitrogen contents. Calcium lacks entirely in the soil. Our field data gave support to the Ellenberg-Landolt ecological indicator values of light and temperature, but not to those of soil reaction and soil nutrient content, for which some changes are proposed.