The Carboniferous coal swamp floras of England: a window on an ancient tropical ecosystem

England has an exceptional range of Westphalian-Stephanian (late Bashkirian-Moscovian) fossil floras spanning some 10 million years. They represent vegetation growing in part of a swamp that covered large areas of tropical Euramerica and which was responsible for the removal of vast quantities of carbon from the atmosphere. This coincided with significant global climatic cooling-the Late Palaeozoic Ice Age. The cratonic Pennine Basin in central and northern England has some of the best preserved fossil floras of this age anywhere in the world, especially notable being those of the Barnsley Thick Seam in Yorkshire and Derbyshire, the Bensham Seam in Northumberland and Durham, and the Coseley Ten Foot Ironstone in the West Midlands. The floras in southern England are mostly not as well preserved but include the historically important Radstock flora of Somerset. The taxonomic diversity dynamics of the fossil floras of the Pennine Basin are rather different from those seen in South Wales, probably due to differences in landscape and habitat, which in turn probably reflect the different tectonic settings. However, evidence of a significant change from lycophyte- to fern-dominated vegetation in latest Westphalian times, recognisable across Euramerica, can be seen in the English floras.