The role of dung beetles as a secondary seed disperser after dispersal by frugivore mammals in a temperate deciduous forest

► We studied the role of dung beetles as secondary seed disperser in temperate forest. ► We used three dung beetle species and bear feces within five plant species and two type markers. ► Most seeds were buried to a 3-6 cm depth, and they were germinated on high rate. ► Burial of the seeds was only observed by P. auratus with large body size. ► Dung beetle is also act as a secondary seed disperser temperate zone.