Extending the rotation period in larch plantations increases canopy heterogeneity and promotes species richness and abundance of native beetles: Implications for the conservation of biodiversity

► The effects of long-rotation in plantations on beetle diversity were studied. ► The beetle diversity, especially phytophages in canopies, increased with long rotation. ► Among ecological variables, naturally regenerated mature trees showed strong positive influences on beetle diversity. ► These regenerated trees contributed to habitat complexity especially in canopies. ► In long-rotation, naturally regenerated mature native trees in plantations are important for conserving native beetle diversity.