Keywords: Arg9; nona-arginine; CHO; Chinese Hamster Ovary; CPP; cell-penetrating peptide; GAG; glycosaminoglycan; GAGneg; glycosaminoglycan deficient CHO cells; GPMV; giant plasma membrane vesicle; HSneg; heparan sulfate deficient CHO cells; pTat; Tat peptide; TP;