A generalization of Hamilton's rule-Love others how much?

► We provide a general methodology to find evolutionarily stable (ES) heritable traits. ► We formulate a generalized version of Hamilton's rule with this methodology. ► Our approach applies to a wide range of pairwise interactions among kin and non-kin. ► We consider heritable behavior rules that adapt actions to the situation at hand. ► ES rules may yield actions that violate Hamilton's rule at the action level.