Transcriptional control of early vein expression of CYCA2; 1 and CYCA2;4 in Arabidopsis leaves

► CYCA2;1 and CYCA2;4 are expressed at preprocambial stages of vein development. ► CYCA2;1 preprocambial expression depends on a 76-bp cis-regulatory element. ► CYCA2;4 preprocambial expression depends on a 77-bp cis-regulatory element. ► Preprocambial expression is encoded in regulatory elements of different structures.