Network models of frugivory and seed dispersal: Challenges and opportunities

► Network analyses are a popular tool to study frugivory and seed dispersal mutualisms. ► We suggest new research avenues to connect network models to ecological processes. ► Using a field study, we empirically tested a network model prediction. ► Model predictions were refuted because the model ignored key behavioral and habitat details. ► Opportunities lie ahead to use network models to generate process-based hypotheses and field tests.