Size variation of conodont elements of the Hindeodus–Isarcicella clade during the Permian–Triassic transition in South China and its implication for mass extinction

Based on the analysis of thousands of conodont specimens from the Permian –Triassic (P–T) transition at Meishan (the GSSP of P–T Boundary) and Shangsi sections in South China, this study investigates the size variation of Hindeodus and Isarcicella P1 elements during the mass extinction interval. The results demonstrate that Hindeodus–Isarcicella underwent 4 episodes of distinct size reduction during the P–T transition at the Meishan Section and 2 episodes of size reduction in the earliest Triassic at Shangsi. The size reductions at Meishan took place at the junctions of beds 24d/24e, 25/26, 27b/27c and 28/29, and at the junctions of beds 28/29c and 30d/31a at Shangsi. The two earliest Triassic size reduction episodes were correlative between the two sections. These changes coincide with some important geological events such as eustatic sea-level changes, anoxic events, carbon isotope oscillations, miniaturization of brachiopods and microbial changes. Through detailed investigation of the palaeoenvironment and the palaeoecology of Hindeodus–Isarcicella, the authors propose that the main causes of the size reduction was a sharp decline of food availability because of the mass extinction and the anoxic event during the P/T transition. The pattern of size reduction supports suggestions that the end-Permian mass extinction was multi-episodal, consisting of 3 extinction events rather than a single catastrophic event.