Ecological succession evidence in an Upper Jurassic coral reef system (Izwarn section, High Atlas, Morocco)

Late Jurassic coral reefs are present in the Western High Atlas of Morocco along the eastern paleomargin of the young Atlantic Ocean. The Izwarn section shows two reef units made of scleractinian corals and microbialites, included in a progradational shallowing-upward succession. Changes in reefal composition and structure observed in a ca. 50 cm-thick interval at the base of the reef unit 1 are interpreted to represent the first well-documented ecological succession in Late Jurassic reefs. Bioclastic deposits, sparse and poorly diversified corals (five genera) preserved in life position, and finally a more diversified coral assemblage (11 genera) building a reef framework with stromatolitic crusts reflect pioneer, colonization and diversification stages, respectively. A specific depth window, where light was reduced but sufficient for a catch-up growth mode and below the erosive action of waves, probably at the lower slope of a fore-reef area, has made possible the development and the record of this ecological succession. With a more diversified coral assemblage (16 genera), the reef unit 2 reflects shallower and well-agitated waters of a fore-reef area. Overlaying this second reef unit, the upper part of the section reveals back-reef deposits made of reworked reef elements, whereas a putative reef crest area is not recorded.