Kernel morphometric traits in hexaploid wheat (Triticum aestivum L.) are modulated by intricate QTL × QTL and genotype × environment interactions

► Fifty nine wheat kernel size and shape QTLs mapped on a linkage map of 251 SSR markers. ► Six stable QTLs could be promising marker assisted selection targets. ► Epistatic, pleiotropic and environmentally modulated QTLs detected.