Intraspecific morphological variation in late Miocene Bolboforma, and implications for their classification, ecology, and biostratigraphic utility

The distribution of bolboforms in (mainly) southern hemisphere marine sediments appears to track the palaeocirculation of Southern Component Intermediate Water (SCIW). Hence addition of calcite to the outer surface of the test wall probably occurred while the bolboforms were entrained in subsurface flows of SCIW. In this respect, the increased complexity of ornamentation may have been a morphological adaptation to counter the loss of buoyancy as calcite was added to the outer surface of the test wall. This would have presumably contributed to the entrainment and dispersal of bolboforms in the flow of subsurface water masses, and it suggests that the tests of some bolboforms at least may represent a dispersal phase in the life cycle of this enigmatic group of microfossils.