Chromatin repositioning activity and transcription machinery are both recruited by Ace1p in yeast CUP1 activation

► We examine how transcriptional activator Ace1p regulates CUP1 induction. ► Ace1p is required for CUP1 transcriptional activation. ► Ace1p directly participates in CUP1 activation. ► Ace1p is required to recruit gene-wide chromatin remodeling activity. ► Ace1p is required to recruit the transcription machinery.