Discovery of the enigmatic genus Skiapus Morley, 1917 (Hymenoptera: Ichneumonidae: Ophioninae) from Japan

The enigmatic ophionine genus Skiapus Morley, 1917 of the family Ichneumonidae is discovered from Japan based on S. niger Sheng, 2006, which had been known from China. We also recognized following two species-groups of Skiapus for the first time: (1) the Afrotropical species-group, including only S. coalescens Morley, 1917, characterized by the mandible twisted at 90° with a single apical tooth; and (2) the Eastern Palaearctic species-group, including S. baeki Lee, 2007 and S. niger, characterized by the mandible twisted at 75° with two apical teeth. A key to the Japanese ophionine genera is also provided.