Scydmobisetia gen. nov., the first definite Glandulariini from Upper Cretaceous Burmese amber (Coleoptera: Staphylinidae: Scydmaeninae)

The Glandulariini (=Cyrtoscydmini) includes about 60 genera comprising over 70% of the extant species of Scydmaeninae. This successful group is today very common and abundant in forests of all continents. However, while extinct representatives of much smaller extant tribes, as Clidicini, Eutheiini and Cephenniini, were described from Cretaceous deposits, named genera and species of Glandulariini were so far known only from much younger Baltic, Sicilian and Dominican ambers. Here we report the first genus and species of definite Glandulariini from the Upper Cretaceous Burmese amber, Scydmobisetia vetutissima Jałoszyński and Yamamoto, gen. and sp. nov. Scydmobisetia is the first known glandulariine beetle with a pair of very long vertexal sensilla chaetica, previously not found in this tribe, but common among unrelated Staphylinidae. The new genus shows similarities to the extant genera Anthicimimus, Sciacharis, Spinosciacharis and Horaeomorphus, but S. vetutissima has protibial modifications typical of males of some extant species of Euconnus. We discuss the apparent disproportion in the oldest fossil record of various tribes of scydmaenines as a result of identification problems, and not the lack of known fossils.