Fine structure of granal thylakoid membrane organization using cryo electron tomography

► Cryo electron tomography of ice-embedded isolated grana membrane stacks. ► Tomographic reconstruction revealed densities of Photosystem II core complex. ► 3D structure of the Photosystem II was solved at 40 Å resolution. ► Novel protein densities were revealed at the lumenal side of the Photosystem II.